1986-1989 University of Istanbul, Dept. of Clinical Biochemistry, Istanbul, Turkiye
1988-1989 University of Vienna, Dept. of Medical Chemistry, Vienna. Austria
1989-1990 University of South Florida, Moffitt Cancer Center, Tampa FL
1991-1992 Beth Israel Medical Center, Dept. of Neurosurgery, NYC, NY
1992-1994 Beth Israel Medical Center, Dept. of Pathology, NYC, NY
1994-1995 Memorial Sloan-Kettering Cancer Center, Dept. of Pathology NYC, NY
1995-1997 State University of New York at Stony Brook, Stony Brook, NY
1997-2002 Johns Hopkins Medical School, Dept. of Pathology, Baltimore, MD.
1999-2002 Johns Hopkins Bayview Medical Center, Dept. Pathology, Baltimore, MD
2002-2024 UCSF School of Medicine, Neuropathology Division
2013-2024 Koc University School of Medicine, Istanbul Turkiye
2024- University of Miami, Miller School of Medicine, Dept of Pathology, Miami FL